The Power That Preserves | My Web Site Page 025 Chapter 02 Page 01

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One of the chief preliminary postulates of sexual selection is the unequal number of individuals in the two sexes, for if every male immediately finds his mate there can be no competition for the possession of the female. Darwin has shown that, for the most part, the inequality between the sexes is due simply to the fact that there are more males than females, and therefore the males must take some pains to secure a mate. But the inequality does not always depend on the numerical preponderance of the males, it is often due to polygamy; for, if one male claims several females, the number of females in proportion to the rest of the males will be reduced. Since it is almost always the males that are the wooers, we must expect to find the occurrence of secondary sexual characters chiefly among them, and to find it especially frequent in polygamous species. And this is actually the case.

In the south the Lucanians also rose against Rome. The extension of the Roman dominion in the south of the peninsula had brought the state into connection with the Greek cities, which at one period were so numerous and powerful as to give to this part of Italy the name of Magna Graecia.[25] Many of these cities had now fallen into decay through internal dissensions and the conquests of the Lucanians and other Sabellian tribes; but Tarentum, originally a Lacedaemonian colony, still maintained her former power and splendor. The Tarentines naturally regarded with extreme jealousy the progress of the Roman arms in the south of Italy, and had secretly instigated the Etruscans and Lucanians to form a new coalition against Rome. But the immediate cause of the war between the Lucanians and Romans was the assistance which the latter had rendered to the Greek city of Thurii. Being attacked by the Lucanians, the Thurians applied to Rome for aid, and the Consul C. Fabricius not only relieved Thurii, but defeated the Lucanians and their allies in several engagements (B.C. 252). Upon the departure of Fabricius a Roman garrison was left in Thurii.

Thus we have in these apparently random forms of the calcereous bodies, complex adaptations in which every little detail as to direction, curve, and pointing is exactly determined. That they have selection-value in their present perfected form is beyond all doubt, since the animals are enabled by means of them to bore rapidly into the ground and so to escape from enemies. We do not know what the initial stages were, but we cannot doubt that the little improvements, which occurred as variations of the originally simple slimy bodies of the Holothurians, were preserved because they already possessed selection-value for the Synaptidae. For such minute microscopic structures whose form is so delicately adapted to the role they have to play in the life of the animal, cannot have arisen suddenly and as a whole, and every new variation of the anchor, that is, in the direction of the development of the two arms, and every curving of the shaft which prevented the tips from projecting at the wrong time, in short, every little adaptation in the modelling of the anchor must have possessed selection-value. And that such minute changes of form fall within the sphere of fluctuating variations, that is to say, _that they occur_ is beyond all doubt.

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