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In cases such as we have discussed, it is obvious that there is no possible explanation except through selection. This brings us to the last kind of secondary sexual characters, and the one in regard to which doubt has been most frequently expressed,--decorative colours and decorative forms, the brilliant plumage of the male pheasant, the humming-birds, and the bird of Paradise, as well as the bright colours of many species of butterfly, from the beautiful blue of our little Lycaenidae to the magnificent azure of the large Morphinae of Brazil. In a great many cases, though not by any means in all, the male butterflies are "more beautiful" than the females, and in the Tropics in particular they shine and glow in the most superb colours. I really see no reason why we should doubt the power of sexual selection, and I myself stand wholly on Darwin's side. Even though we certainly cannot assume that the females exercise a conscious choice of the "handsomest" mate, and deliberate like the judges in a court of justice over the perfections of their wooers, we have no reason to doubt that distinctive forms (decorative feathers), and colours have a particularly exciting effect upon the female, just as certain odours have among animals of so many different groups, including the butterflies. The doubts which existed for a considerable time, as a result of fallacious experiments, as to whether the colours of flowers really had any influence in attracting butterflies have now been set at rest through a series of more careful investigations; we now know that the colours of flowers are there on account of the butterflies, as Sprengel first showed, and that the blossoms of Phanerogams are selected in relation to them, as Darwin pointed out.

Besides the three old masters, as we may call them, Buffon, Erasmus Darwin, and Lamarck, there were other quite convinced pre-Darwinian evolutionists. The historian of the theory of descent must take account of Treviranus whose _Biology or Philosophy of Animate Nature_ is full of evolutionary suggestions; of Etienne Geoffroy St. Hilaire, who in 1830, before the French Academy of Sciences, fought with Cuvier, the fellow-worker of his youth, an intellectual duel on the question of descent; of Goethe, one of the founders of morphology and the greatest poet of Evolution--who, in his eighty-first year, heard the tidings of Geoffrey St. Hilaire's defeat with an interest which transcended the political anxieties of the time; and of many others who had gained with more or less confidence and clearness a new outlook on Nature. It will be remembered that Darwin refers to thirty-four more or less evolutionist authors in his Historical Sketch, and the list might be added to. Especially when we come near to 1858 do the numbers increase, and one of the most remarkable, as also most independent champions of the evolution-idea before that date was Herbert Spencer, who not only marshalled the arguments in a very forcible way in 1852, but applied the formula in detail in his _Principles of Psychology_ in 1855.[13]

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